Summary
            Every stage of tree growth may be attacked by one or more species of mammal. Often a species may cause damage at several growth stages. Most mammal damage to trees is from either:
- Browsing – feeding on buds, shoots and foliage
 
- Bark stripping from main stems or branches – gnawing or rubbing.
 
When trying to identify the cause of damage, the most important things to look for are:
- Form of damage (i.e. browsing, gnawing or rubbing)
 
- Height of damage
 
- Time of year when damage occurred
 
- Presence and size of teeth marks
 
- Signs of animal presence and abundance – droppings, footprints, runs, scrapes or burrows.
 
Points to note:
- Lack of teeth in front upper jaw of all deer species produces ragged edge on damaged twigs
 
- The teeth of rabbits and hares produce a sharp knife like cut. Muntjac may bite partly through thin tall stems and pull them down to eat
 
- Sheep and deer browsing damage is often very similar in form, but sheep tend to leave wool evidence
 
- Fraying is a rubbing injury caused when male deer rub new antlers to remove ‘velvet’ or to mark territories.
 
           
        
        
                  
            Findings and Recommendations
            The following three tables show the main characteristics of damage by:
- wild deer
 
- other wild mammals including rabbits, hares, squirrels, mice, voles, edible dormice, moles and badgers
 
- domestic livestock
 
Brackets in the ‘age of trees’ column denote damage is uncommon.
Table A: Wild deer
| Species | 
Age of trees affected | 
Typical signs of damage to trees | 
Comments; damage to other parts of woodland etc. | 
| Red deer | 
- Establishment
 
- Thicket
 
- Pole stage
 
- Mature
 
 
 | 
- Browse seedlings to thicket stage
 
- Fraying and rubbing on bark up to 1.8 m
 
 
 | 
- Damage to poor fences
 
- Severe damage to herb layer
 
 
 | 
| Sika deer | 
- Establishment
 
- Thicket
 
- Pole stage
 
- Mature
 
 
 | 
- Browse seedlings to thicket stage
 
- Scoring of pole stage trunks
 
- Fraying and rubbing on bark up to 1.8 m
 
 
 | 
 | 
| Fallow deer | 
- Establishment
 
- Thicket
 
- Pole stage
 
- (Mature)
 
 
 | 
- Browse seedlings to thicket stage
 
- Fraying bark up to 1.5 m
 
- May pull up recently planted trees
 
 
 | 
- Severe damage to herb layer
 
 
 | 
| Roe deer | 
- Establishment
 
- Thicket
 
- Pole stage
 
- Mature
 
 
 | 
- Browse seedlings to thicket stage
 
- Fraying bark up to 1.2 m
 
 
 | 
- Severe damage to herb layer
 
 
 | 
| Muntjac deer | 
 | 
- Browse seedlings to thicket stage
 
- Fraying bark up to 1 m
 
- May partly bite through taller stems and pull down to browse
 
 
 | 
- Capable of severe damage to herb layer
 
 
 | 
 
Table B: Other wild mammals
| Species | 
Age of trees affected | 
Typical signs of damage to trees | 
Comments; damage to other parts of woodland etc. | 
| Voles (bank & field) | 
 | 
- Eating seeds, seedlings, root cutting of young planted stock
 
- Ringbarking up to 10 cm
 
- Teethmarks only 2 mm wide
 
- Bank vole will climb saplings and eat bark around base of branch
 
 
 | 
- Typical runways in grass with dropping and cut grass piles evident
 
- Nests in tree shelters
 
 
 | 
| Mice (wood, yellow-necked & house) | 
 | 
 | 
- Seed stores
 
- Nests in tree shelters or under mulch mats
 
 
 | 
| Grey squirrel | 
- Seedling
 
- Thicket
 
- Pole stage
 
- Mature
 
 
 | 
- Stripping bark anywhere on stem or branches of pole stage and mature trees
 
- Bark stripping at bottom 50 cm of stem may be confused with rabbit damage
 
 
 | 
- Eating larger seeds
 
- Predation of bird nests
 
 
 | 
| Edible dormouse | 
 | 
- Spiral bark stripping at branch bases
 
 
 | 
- Very restricted range
 
- Hibernates below ground
 
- Compete for tree holes
 
 
 | 
| Rabbit | 
- Establishment
 
- Seedling
 
- Thicket
 
- Pole stage
 
- (Mature)
 
 
 | 
- Cutting stems of planted saplings
 
- Ringbarking bottom 50 cm of stem
 
- Burrows assist windblow
 
- Sharp angled knife-like cuts on end of small stems or branches
 
- Removed portion often eaten
 
 
 | 
- Most vegetation in area around burrow often grazed very low
 
 
 | 
| Hare (mountain & brown) | 
 | 
- Often sporadic but widespread
 
- May eat along a row of young trees
 
- Damage up to 70 cm
 
 
 | 
 | 
| Badger | 
 | 
- Setts under roots
 
- Limited bark damage
 
 
 | 
- Create holes under fences for other pests to gain entry
 
 
 | 
| Mole | 
 | 
- Tunnelling may cause desiccation of seedlings and transplant roots
 
- Soil heaps may bury young plants
 
 
 | 
- Assists drainage of gley soils
 
 
 | 
 
Table C: Domestic livestock
| Species | 
Age of trees affected | 
Tree and other collateral damage | 
| Sheep | 
- Establishment
 
- Thicket
 
- (Pole stage)
 
 
 | 
- Removal of ground vegetation
 
- Browsing and bark stripping
 
- Newly planted trees may be pulled out
 
 
 | 
| Goat | 
- Establishment
 
- Thicket
 
- Pole stage
 
 
 | 
- Removal of ground vegetation
 
- Browsing and bark stripping
 
- Newly planted trees may be pulled out
 
 
 | 
| Cattle | 
- Establishment
 
- Thicket
 
- Pole stage
 
 
 | 
- Removal of ground vegetation and newly planted trees or natural regeneration
 
- Treading impacts may be detrimental to roots or beneficial by providing nutrient and germination patches
 
- Coarse browsing of foliage to 1.5 m
 
- Newly planted trees may be pulled out
 
 
 | 
| Pigs & feral boar | 
- Seedling
 
- Establishment
 
- Thicket
 
- Pole stage
 
- (Mature)
 
 
 | 
- Removal of large seeds, ground vegetation and natural regeneration
 
- Browsing and root damage by grubbing
 
- Digging rabbit burrows, holes created under fences
 
- Rubbing on trunks
 
 
 | 
| Ponies | 
- Establishment
 
- Thicket
 
- (Pole stage)
 
 
 | 
- Newly planted trees may be pulled out
 
- Browsing to 2 metres
 
- Bark stripping with characteristic diagonal teeth marks from both jaws
 
- Grazing shrubs and ground flora
 
 
 |